Two of the four cubs of a tigress shifted from Corbett Tiger Reserve have been killed by leopards in the Berivada range of Rajaji Tiger Reserve (RTR), officials said on Tuesday.

RTR director Saket Badola said one of the cubs was male and the other female.

He told that it is estimated that the cubs will be about one to one and a half months old.

One cub's neck was broken and the other's head was crushed, Badola said.

The transferred tigress had given birth to four cubs in the Chillawali range on May 24.

Now only two cubs of this tigress are left alive.